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Bald Eagle
Eagle
Names
Common Name: Bald Eagle
Scientific Name: Haliaeetus leucocephalus
Appearance
The plumage of an adult Bald Eagle is evenly dark brown with a white head and tail. The tail is moderately long and slightly wedge-shaped. Males and females are identical in plumage coloration, but sexual dimorphism is evident in the species in that females are generally 25% larger than males
Behavior
very teritorrial
Please note that this information is taken from the series and may not be factual.

The Bald Eagle (Haliaeetus leucocephalus) is a bird of prey found in North America. A sea eagle, it has two known sub-species and forms a species pair with the White-tailed Eagle (Haliaeetus albicilla). Its range includes most of Canada and Alaska, all of the contiguous United States, and northern Mexico. It is found near large bodies of open water with an abundant food supply and old-growth trees for nesting.

The Bald Eagle is an opportunistic feeder which subsists mainly on fish, which it swoops down and snatches from the water with its talons. It builds the largest nest of any North American bird and the largest tree nests ever recorded for any animal species, up to 4 meters (13 ft) deep, 2.5 meters (8.2 ft) wide, and one metric ton (1.1 tons) in weight.Sexual maturity is attained at the age of four to five years.

Bald Eagles are not actually bald; the name derives from an older meaning of "white headed". The adult is mainly brown with a white head and tail. The sexes are identical in plumage, but females are generally larger than males with a deeper beak [1]. The beak is large and hooked. The plumage of the immature is brown.

The Bald Eagle is the national bird of the United States of America and appears on its Seal. In the late 20th century it was on the brink of extirpation in the continental United States. Populations recovered and the species was removed from the U.S. federal government's list of endangered species on July 12, 1995 and transferred to the list of threatened species. It was removed from the List of Endangered and Threatened Wildlife in the Lower 48 States on June 28, 2007.

Description

The plumage of an adult Bald Eagle is evenly dark brown with a white head and tail. The tail is moderately long and slightly wedge-shaped. Males and females are identical in plumage coloration, but sexual dimorphism is evident in the species in that females are generally 25% larger than males.[2] The beak, feet and irides are bright yellow. The legs are feather-free, and the toes are short and powerful with large talons. The highly developed talon of the hind toe is used to pierce the vital areas of prey while it is held immobile by the front toes.[3] The beak is large and hooked, with a yellow cere.[4] The adult Bald Eagle is unmistakable in its native range. The closely related African Fish Eagle (H. vocifer) (from far outside of the Bald Eagle's range) also has a brown body, white head and tail, but differs from the Bald in having a white chest and black tip to the bill.[5]

The plumage of the immature is a dark brown overlaid with messy white streaking until the fifth (rarely fourth, very rarely third) year, when it reaches sexual maturity.[2][3] Immature Bald Eagles are distinguishable from the Golden Eagle (Aquila chrysaetos), the only other very large, non-vulturine bird in North America, in that the former has a larger, more protruding head with a larger beak, straighter edged wings which are held flat (not slightly raised) and with a stiffer wing beat and feathers which do not completely cover the legs. When seen well, the Golden Eagle is distinctive in plumage with a more solid warm brown color than an immature Bald Eagle, with a reddish-golden patch to its nape and (in immature birds) a highly contrasting set of white squares on the wing.[6]

The Bald Eagle has sometimes been considered the largest true raptor (accipitrid) in North America. The only larger species of raptor-like bird is the California Condor (Gymnogyps californianus), a New World vulture which today are not generally considered a taxonomic ally of true accipitrids.[7] However, the Golden Eagle, averaging 4.18 kg (9.2 lb) and 63 cm (25 in) in wing chord length in its American race (A. c. canadensis), is merely 455 g (1.00 lb) lighter in mean body mass and exceeds the Bald Eagle in mean wing chord length by around 3 cm (1.2 in).[5][8] Additionally, the Bald Eagle's close cousins, the relatively longer-winged but shorter-tailed White-tailed Eagle and the overall larger Steller's Sea Eagle (H. pelagicus), may rarely vagrate to coastal Alaska from Asia.[5]

The Bald Eagle has a body length of 70–102 centimeters (28–40 in). Typical wingspan is between 1.8 and 2.3 m (5.9 and 7.5 ft) and mass is normally between 3 and 6.3 kilograms (6.6 and 14 lb).[5] Females are about 25% larger than males, averaging 5.6 kg (12 lb), and against the males' average weight of 4.1 kg (9.0 lb).[2][9][10][11] The size of the bird varies by location and generally corresponds with Bergmann's rule, since the species increases in size further away from the Equator and the tropics. The smallest specimens are those from Florida, where mature males may weigh as little as 2.3 kg (5.1 lb) and have a wingspan of 1.68 m (5.5 ft). Similarly small, eagles from South Carolina average 3.27 kg (7.2 lb) in mass and 1.88 m (6.2 ft) in wingspan.[12] The largest eagles are from Alaska, where large females may weigh up to 7.5 kg (17 lb) and span 2.44 m (8.0 ft) across the wings.[4][13] A survey of adult weights in Alaska showed that females weighed on average 6.3 kg (14 lb) and males weighed 4.3 kg (9.5 lb).[14] Among standard linear measurements, the wing chord is 51.5–69 cm (20.3–27 in), the tail is 23–37 cm (9.1–15 in) long, and the tarsus is 8 to 11 cm (3.1 to 4.3 in).[5][15] The culmen reportedly ranges from 3 to 7.5 cm (1.2 to 3.0 in), while the measurement from the gape to the tip of the bill is 7–9 cm (2.8–3.5 in).[15][16]

The call consists of weak staccato, chirping whistles, kleek kik ik ik ik, somewhat similar in cadance to a gull's call. The calls of young birds tend to be more harsh and shrill than those of adults.[5][6]

Taxonomy

The Bald Eagle placed in the genus Haliaeetus (sea eagles) which gets both its common and specific scientific names from the distinctive appearance of the adult's head. Bald in the English name is derived from the word piebald, and refers to the white head and tail feathers and their contrast with the darker body.[17] The scientific name is derived from Haliaeetus, New Latin for "sea eagle" (from the Ancient Greek haliaetos), and leucocephalus, Latinized Ancient Greek for "white head," from λευκος leukos ("white") and κεφαλη kephale ("head").[18][19]

The Bald Eagle was one of the many species originally described by Linnaeus in his 18th century work Systema Naturae, under the name Falco leucocephalus.[20]

There are two recognized subspecies of Bald Eagle:[2][21]

  • H. l. leucocephalus (Linnaeus, 1766) is the nominate subspecies. It is separated from H. l. washingtoniensis at approximately latitude 38° N, or roughly the latitude of San Francisco.[22] It is found in the southern United States and Baja California.[23]
  • H. l. washingtoniensis (Audubon, 1827), synonym H. l. alascanus Townsend, 1897, the northern subspecies, is larger than southern nominate leucocephalus. It is found in the northern United States, Canada and Alaska.[2][23] This subspecies reaches further south than latitude 38° N on the Atlantic Coast, where they occur in the Cape Hatteras area.[22]

The Bald Eagle forms a species pair with the Eurasian White-tailed Eagle. This species pair consists of a white-headed and a tan-headed species of roughly equal size; the White-tailed Eagle also has overall somewhat paler brown body plumage. The two species fill the same Ecological niche in their respective ranges. The pair diverged from other Sea Eagles at the beginning of the Early Miocene (c. 10 Ma BP) at the latest, but possibly as early as the Early/Middle Oligocene, 28 Ma BP, if the most ancient fossil record is correctly assigned to this genus.[24] The two species probably diverged in the North Pacific, as the White-tailed Eagle spread westwards into Eurasia and the Bald Eagle spread eastwards into North America.[25]

Range

The Bald Eagle's natural range covers most of North America, including most of Canada, all of the continental United States, and northern Mexico. It is the only sea eagle endemic to North America. Occupying varied habitats from the bayous of Louisiana to the Sonoran Desert and the eastern deciduous forests of Quebec and New England, northern birds are migratory, while southern birds are resident, remaining on their breeding territory all year. At minimum population, in the 1950s, it was largely restricted to Alaska, the Aleutian Islands, northern and eastern Canada, and Florida.[26] Today, they are much more common (almost attaining their peak numbers pre-colonization in North America), and nest in every continental state and province in the United States and Canada.[27]

Bald Eagles will also congregate in certain locations in winter. From November until February, one to two thousand birds winter in Squamish, British Columbia, about halfway between Vancouver and Whistler. The birds primarily gather along the Squamish and Cheakamus Rivers, attracted by the salmon spawning in the area.[28]

It has occurred as a vagrant twice in Ireland; a juvenile was shot illegally in Fermanagh on January 11, 1973 (misidentified at first as a White-tailed Eagle), and an exhausted juvenile was captured in Kerry on November 15, 1987.[29]

Habitat

[2][3]Juvenile with salmon, Katmai National Park[4][5]With freshly caught fish in KodiakThe Bald Eagle occurs during its breeding season in virtually any kind of American wetland habitat such as seacoasts, rivers, large lakes or marshes or other large bodies of open water with an abundance of fish. Studies have shown a preference for bodies of water with a circumference greater than 11 km (7 mi), and lakes with an area greater than 10 square kilometers (4 sq mi) are optimal for breeding Bald Eagles.[30]

The Bald Eagle typically requires old-growth and mature stands of coniferous or hardwood trees for perching, roosting, and nesting. Tree species reportedly is less important to the eagle pair than the tree's height, composition and location.[31] Perhaps of paramount importance for this species is an abundance of comparatively large trees surrounding the body of water. Selected trees must have good visibility, be over 20 m (66 ft) tall, an open structure, and proximity to prey. If nesting trees are in standing water such as in a mangrove swamp, the nest can be located fairly low, at as low 6 m (20 ft) above the ground.[32] In a more typical tree standing on dry ground, nests may be located from 16 to 38 m (52 to 125 ft) in height. In Chesapeake Bay, nesting trees averaged 82 cm (32 in) in diameter and 28 m (92 ft) in total height, while in Florida, the average nesting tree stands 23 m (75 ft) high and is 23 cm (9.1 in) in diameter.[33][34] Trees used for nesting in the Greater Yellowstone area average 27 m (89 ft) high.[35] Trees or forest used for nesting should have a canopy cover of no more than 60%, and no less than 20%, and be in close proximity to water.[30] Most nests have been found within 200 m (660 ft) of open water. The greatest distance from open water recorded for a Bald Eagle nest was over 3 km (1.9 mi), in Florida.[7]

In Florida, nesting habitats often consist of mangrove swamps, the shorelines of lakes and rivers, pinelands, seasonally flooded flatwoods, hardwood swamps, and open prairies and pastureland with scattered tall trees. Favored nesting trees in Florida are Slash Pines (Pinus elliottii), Longleaf Pines (P. palustris), Loblolly Pines (P. taeda) and cypress trees, but for the southern coastal areas where mangroves are usually used.[32] In Wyoming, groves of mature cottonwoods or tall pines found along streams and rivers are typical bald eagle nesting habitats. Wyoming eagles may inhabit habitat types ranging from large, old-growth stands of Ponderosa Pines (Pinus ponderosa) to narrow strips of riparian trees surrounded by rangeland.[7] In Southeast Alaska, Sitka Spruce (Picea sitchensis) provided 78% of the nesting trees used by eagles, followed by hemlocks (Tsuga) at 20%.[31] Increasingly, eagles nest in man-made reservoirs stocked with fish.[32]

The Bald Eagle is usually quite sensitive to human activity while nesting, and is found most commonly in areas with minimal human disturbance. It chooses sites more than 1.2 km (0.75 mi) from low-density human disturbance and more than 1.8 km (1.1 mi) from medium- to high-density human disturbance.[30] However, Bald Eagles will occasionally venture into large estuaries or secluded groves within major cities, such as Hardtack Island on the Willamette River in Portland, Oregon or John Heinz National Wildlife Refuge at Tinicum in Philadelphia, Pennsylvania, which are surrounded by a great quantity of human activity.[36][37] Even more contrary to the usual sensitivity to disturbance, a family of Bald Eagles moved to the Harlem neighborhood in New York City in 2010.[38]

While wintering, Bald Eagles tend to be less habitat and disturbance sensitive. They will commonly congregate at spots with plentiful perches and waters with plentiful prey and (in Northern climes) partially unfrozen waters. Alternately, non-breeding or wintering Bald Eagles, particularly in areas with a lack of human disturbance, spend their time in various upland, terrestrial habitats sometimes quite far away from waterways. In the Northern half of North America (especially the interior portion), this terrestrial inhabitance by Bald Eagles tends to be especially prevalent because unfrozen water may not be accessible. Upland wintering habitats often consist of open habitats with concentrations of medium-sized mammals, such as prairies, meadows or tundra, or open forests with regular carrion access.[7][31]

Behavior

[6][7]Head detailsThe Bald Eagle is a powerful flier, and soars on thermal convection currents. It reaches speeds of 56–70 kilometers per hour (35–43 mph) when gliding and flapping, and about 48 kilometers per hour (30 mph) while carrying fish.[39] Its dive speed is between 120–160 kilometers per hour (75–99 mph), though it seldom dives vertically.[40] It is partially migratory, depending on location. If its territory has access to open water, it remains there year-round, but if the body of water freezes during the winter, making it impossible to obtain food, it migrates to the south or to the coast. A number of populations are subject to post-breeding dispersal, mainly in juveniles; Florida eagles, for example, will disperse northwards in the summer.[41] The Bald Eagle selects migration routes which take advantage of thermals, updrafts, and food resources. During migration, it may ascend in a thermal and then glide down, or may ascend in updrafts created by the wind against a cliff or other terrain. Migration generally takes place during the daytime, usually between the local hours of 8:00 a.m. and 6:00 p.m., when thermals are produced by the sun.[3]

Diet and feeding

The Bald Eagle is an opportunistic carnivore with the capacity to predate a great variety of prey. Throughout their range, fish often compromise the lion's share of the eagle's diet.[42] In 20 food habit studies across the species' range, fish comprised 56% of the diet of nesting eagles, birds 28%, mammals 14% and other prey 2%.[43] In Southeast Alaska, fish comprise approximately 66% of the year-around diet of Bald Eagles and 78% of the prey brought to the nest by the parents.[44] Eagles living in the Columbia River Estuary in Oregon were found to rely on fish for 90% of their dietary intake.[45] In the Pacific Northwest, spawning trout and salmon provide most of the Bald Eagles' diet from late summer throughout fall.[46] Southeast Alaskan eagles largely predate Pink salmon (Oncorhynchus gorbuscha), Coho salmon (O. kisutch) and, more locally, Sockeye salmon (O. nerka), with Chinook salmon (O. tshawytscha), due to their large size (12 to 18 kg (26 to 40 lb) average adult size) probably being taken only as carrion.[44] Also important in the estuaries and shallow coastlines of southern Alaska are Pacific herring (Clupea pallasii), Pacific sand lance (Ammodytes hexapterus) and Eulachon (Thaleichthys pacificus).[44] In Oregon's Columbia River Estuary, the most significant prey species were Largescale suckers (Catostomus macrocheilus) (17.3% of the prey selected there), American shad (Alosa sapidissima; 13%) and Common Carp (Cyprinus carpio; 10.8%).[45] Eagles living in the Chesapeake Bay in Maryland were found to subsist largely on American gizzard shad (Dorosoma cepedianum), Threadfin shad (D. petenense) and White bass (Morone chrysops).[47] Floridian eagles have been reported to predate catfish, mostly prevalently the Brown bullhead (Ameiurus nebulosus) and any species in the Ictalurus genus as well as mullet, trout, needlefish, and eels.[7][32][48] Wintering eagles on the Platte River in Nebraska preyed mainly on American gizzard shads and Common Carp.[49] From observation in the Columbia River, 58% of the fish were caught directly by the predating eagle, 24% were scavenged as carcasses and 18% were pirated away from other animals.[45]

Even eagles living in relatively arid regions still typically rely primarily on fish as prey. In Sonora, Mexico and Arizona, 77% and over 73%, respectively, of prey remains at the nests were from fish, largely various catfish and Rainbow trout (Oncorhynchus mykiss). Prey fish targeted by Bald Eagles are often quite large. When experimenters offered fish of different sizes in the breeding season around Lake Britton in California, fish measuring 34 to 38 cm (13 to 15 in) were taken 71.8% of the time by parent eagles while fish measuring 23 to 27.5 cm (9.1 to 10.8 in) were chosen only 25% of the time.[50] At nests around Lake Superior, the remains of fish (mostly suckers) were found to average 35.4 cm (13.9 in) in total length.[51] In the Columbia River estuary, most predated by eagles were estimated to measure between 30 and 60 cm (12 and 24 in) in length, and carp flown with (laboriously) were up to 86 cm (34 in) in length.[45]

Benthic fishes such as catfish are usually consumed after they die and float to the surface, though while temporarily swimming in the open may be more vulnerable to predation than most fish since their eyes focus downwards.[47] Bald Eagles also regularly exploit water turbines which produce battered, stunned or dead fish easily consumed.[52] Predators who leave behind scraps of dead fish that they kill, such as Brown Bears (Ursus arctos), Gray Wolves (Canis lupus) and Red Foxes (Vulpes vulpes), may be habitually followed in order to scavenge the kills secondarily.[44] Once North Pacific salmon die off after spawning, usually local Bald Eagles eat salmon carcasses almost exclusively. Eagles in Washington need to consume 489 g (1.08 lb) of fish each day for survival, with adults generally consuming more than juveniles and thus reducing potential energy deficiency and increasing survival during winter.[43]

Behind fish, the next most significant prey base for Bald Eagles are other waterbirds. The contribution of such birds to the eagle's diet is variable, depending on the quantity and availability of fish near the water's surface. Waterbirds can seasonally comprise from 7% to 80% of the prey selection for eagles in certain localities.[45][53] Exceptionally, in the Greater Yellowstone area, birds were eaten as regularly as fish year-around, with both prey groups comprising 43% of the studied dietary intake.[35] Preferred avian prey includes grebes, alcids, ducks, gulls, coots, herons, egrets, and geese.[54] Bird species most preferred as prey by eagles tend to be medium-sized, such as Western Grebes (Aechmophorus occidentalis), Mallards (Anas platyrhynchos) and American coots (Fulica americana) as such prey is relatively easy for the much larger eagles to catch and fly with.[7][45] American Herring Gull (Larus smithsonianus) are the favored avian prey species for eagles living around Lake Superior.[51] Larger waterbirds are occasionally predated as well, with wintering Emperor Geese (Chen canagica) and Snow Geese (C. caerulescens), which gather in large groups, sometimes becoming regular prey.[15][55] Other large waterbirds hunted at least occasionally by Bald Eagles have included Common Loons (Gavis immer),[56] Great Black-backed Gulls (Larus marinus),[57] Sandhill Cranes (Grus canadensis),[58] Great Blue Herons (Ardea herodias),[43] Canada Geese (Branta canadensis),[47] Brown Pelicans (Pelecanus occidentalis),[32] and fledging American White Pelicans (P. erythrorhynchos).[59] Colony nesting seabirds, such as alcids, storm petrels, cormorants, Northern Gannets (Morus bassanus), terns and gulls, may be especially vulnerable to predation. Due to easy accessibility and lack of formidable nest defense by such species, Bald Eagles are capable of predating such seabirds at all ages, from eggs to mature adults, and can effectively cull large portions of a colony.[60]

Along some portions of the North Pacific coastline, Bald Eagles which had historically predated mainly kelp-dwelling fish and supplementally Sea Otter (Enhydra lutris) pups are now preying mainly on seabird colonies since both the fish (possibly due to overfishing) and otters (cause unknown) have had precipitious population declines, causing concern for seabird conservation.[61] Because of this more extensive predation, some biologist have expressed concern that murres are heading for a "conservation collision" due to heavy eagle predation.[60] Eagles have been confirmed to attack nocturnally active, burrow-nesting seabird species such as storm petrels and shearwaters by digging out their burrows and feeding on all animals they find inside.[62] If a Bald Eagle flies close by, waterbirds will often fly away en masse, though in other cases they may seemingly ignore a perched eagle. If the said birds are on a colony, this exposed their unprotected eggs and nestlings to scavengers such as gulls.[60] Bird prey may occasionally be attacked in flight, with prey up to the size of Canada Geese attacked and killed in mid-air.[54] Unprecedented photographs of a Bald Eagle unsuccessfully attempting to predate a much larger adult Trumpeter Swan (Cygnus buccinator) in mid-flight were taken recently.[63] While adults often actively predate waterbirds, congregated wintering waterfowl are frequently exploited for carcasses to scavenge by immature eagles in harsh winter weather.[64] Bald Eagles have been recorded as killing other raptors on occasion. In some cases, these may be attacks of competition or kleptoparasitism on rival species but ended with the consumption of the victim. Raptorial birds reported to have be hunted by these eagles have included large adults of species such as Great Horned Owls (Bubo virginianus),[65] Red-tailed Hawks (Buteo jamaicensis),[66] Ospreys (Pandion haliaetus)[67] and Black (Coragyps atratus) and Turkey Vultures (Cathartes aura).[68]

Mammalian prey includes rabbits, hares, ground squirrels, Raccoons (Procyon lotor), Muskrats (Ondatra zibethicus), beavers (Castor canadensis), and deer fawns. Newborn, dead, sickly or already injured mammals are often targeted. However, more formidable prey such as adult raccoons and sub-adult beavers are sometimes attacked. In the Chesapeake Bay area, Bald Eagles are reportedly the main natural predators of raccoons.[69][70] Where available, seal colonies can provide much food. On Protection Island, Washington, commonly feed on Harbor seal (Phoca vitulina) afterbirths, still-borns and sickly seal pups.[71] On San Juan Island in Washington, introduced European rabbits (Oryctolagus cuniculus), mainly those killed by auto accidents, comprise nearly 60% of the dietary intake of eagles.[72] In landlocked areas of North America, wintering Bald Eagles may become habitual predators of medium-sized mammals that occur in colonies or local concentrations, such prairie dogs (Cynomys) and jackrabbits (Lepus).[7][73] Together with the Golden Eagle, Bald Eagles are occasionally accused of predating livestock, especially sheep (Ovis aries). There are a handful of proven cases of lamb predation, some of specimens weighing up to 11 kg (24 lb), by Bald Eagles but they are much less likely to attack a healthy lamb than a Golden Eagle and both species prefer native, wild prey and are unlikely to cause any extensive determent to human livelihoods.[74] There is one case of a Bald Eagle killing and feeding on an adult, pregnant ewe (then joined in eating the kill by at least 3 other eagles), which, weighing on average over 60 kg (130 lb), is much larger than any other known prey taken by this species.[75]

Supplemental prey are readily taken given the opportunity. In some areas reptiles may become regular prey, especially warm areas such as Florida where reptile diversity is high. Turtles are perhaps the most regularly hunted type of reptile.[7] In coastal New Jersey, 14 of 20 studied eagle nests included remains of turtles. The main species found were common musk turtles (Sternotherus odoratus), Diamondback terrapin (Malaclemys terrapin) and juvenile common snapping turtles (Chelydra serpentina). In these New Jersey nests, mainly subadult and small adults were taken, ranging in carapace length from 9.2 to 17.1 cm (3.6 to 6.7 in).[76] Snakes are also taken occasionally, especially partially aquatic ones, as are amphibians and crustaceans (largely crayfish and crabs).[32][45]

To hunt fish, the eagle swoops down over the water and snatches the fish out of the water with its talons. They eat by holding the fish in one claw and tearing the flesh with the other. Eagles have structures on their toes called spicules that allow them to grasp fish. Osprey also have this adaptation.[39] Bald Eagles have powerful talons and have been recorded flying with a 6.8 kg (15 lb) Mule Deer (Odocoileus hemionus) fawn.[77] This feat is the record for the heaviest load carrying ever verified for a flying bird.[78] It has been estimated that the gripping power (pounds by square inch) of the bald eagle is ten times greater than that of a human.[79] Bald eagles can fly with fish at least equal to their own weight, but if the fish is too heavy to lift, the eagle may be dragged into the water. It may swim to safety, but some eagles drown or succumb to hypothermia. Many sources claim that Bald Eagles, like all large eagles, cannot normally take flight carrying prey more than half of their own weight unless aided by favorable wind conditions.[32][55] On numerous occasions, when large prey such as mature salmon or geese are attacked, eagles have been seen to make contact and then drag the prey in a strenuously labored, low flight over the water to a bank, where they then finish off and dismember the prey.[15] When food is abundant, an eagle can gorge itself by storing up to 1 kg (2.2 lb) of food in a pouch in the throat called a crop. Gorging allows the bird to fast for several days if food becomes unavailable.[32] Occasionally, Bald Eagles may hunt cooperatively when confronting prey, especially relatively large prey such as jackrabbits or herons, with one bird distracting potential prey, while the other comes behind it in order to ambush it.[4][80][81] While hunting waterfowl, Bald Eagles repeatedly fly at a target and cause it to dive repeatedly, hoping to exhaust the victim so it can be caught (White-tailed Eagles have been recorded hunting waterfowl in the same way). When hunting concentrated prey, a successful catch which often results in the hunting eagle being pursued by other eagles and needing to find an isolated perch for consumption if it is able to carry it away successfully.[15]

Unlike some other eagle species, Bald Eagles rarely take on evasive or dangerous prey on their own. The species mainly target prey which is much smaller than themselves, with most live fish caught weighing 1 to 3 kg (2.2 to 6.6 lb) and most waterbirds predated weighing 0.2 to 2.7 kg (0.44 to 6.0 lb).[44][55][82] They attain much their food as carrion or via a practice known as kleptoparasitism, where they steal prey away from other predators. Due to their dietary habits, Bald Eagles are frequently viewed in a negative light by humans.[7] Thanks to their superior foraging ability and experience, adults are generally more likely to hunt live prey than immature eagles, which often obtain their food from scavenging.[83][84] They are not very selective about the condition or origin, whether provided by humans, other animals, auto accidents or natural causes, of a carcass's presence, but will avoid eating carrion where disturbances from humans are a regular occurrence. They will scavenge carcasses up to the size of whales, though carcasses of ungulates and large fish are seemingly preferred.[15] Bald Eagles also may sometimes feed on subsistence scavenged or stolen from campsites and picnics, as well as garbage dumps (dump usage is habitual mainly in Alaska).[85]

When competing for food, eagles will usually dominate other fish-eaters and scavengers, aggressively displacing mammals such as Coyotes (Canis latrans) and foxes, and birds such as corvids, gulls, vultures and other raptors.[85] Occasionally, coyotes, Bobcats (Lynx rufus) and domestic dogs (Canis lupus familiaris) can displace eagles from carrion, usually less confident immature birds, as has been recorded in Maine.[86] Bald Eagles are less active, bold predators than Golden Eagles and get relatively more of their food as carrion and from kleptoparasitism (although it is now generally thought that Golden Eagles eat more carrion than was previously assumed).[8] However, the two species are roughly equal in size, aggressiveness and physical strength and so competitions can go either way. Neither species is known to be dominant, and the outcome depends on the size and disposition of the individual eagles involved.[15] The Bald Eagle is thought to be much more numerous in North America than the Golden Eagle, with the Bald species estimated to number at least 150,000 individuals, about twice as many Golden Eagles there are estimated to live in North America.[8][27] Due to this, Bald Eagles often outnumber Golden Eagles at attractive food sources.[8] Despite the potential for contention between these animals, in New Jersey during winter, a Golden Eagle and numerous Bald Eagles were observed to hunt Snow Geese alongside each other without conflict.[87] Similarly, both eagle species have been recorded, via video-monitoring, to feed on gut pills and carcasses of White-tailed Deer (Odocoileus virginianus) in remote forest clearings in the eastern Appalachian Mountains without apparent conflict.[8] Many Bald Eagles are habitual kleptoparasites, especially in winters when fish are harder to come by. They have been recorded stealing fish from other predators such as Ospreys, herons and even otters.[15][88] They have also been recorded opportunistically pirating birds from Peregrine Falcons (Falco peregrinus), prairie dogs from Ferruginous Hawks (Buteo regalis) and even jackrabbits from Golden Eagles.[89][90] When they approach scavengers like dogs, gulls or vultures at carrion sites, they often aggressively attack them and try to force them to disgorge their food.[32] Healthy adult Bald Eagles are not preyed on in the wild and are thus considered apex predators.[91]

Reproduction

[8][9]MatingBald Eagles are sexually mature at four or five years of age. When they are old enough to breed, they often return to the area where they were born. It is thought that Bald Eagles mate for life. However, if one member of a pair dies or disappears, the other will choose a new mate. A pair which has repeatedly failed in breeding attempts may split and look for new mates.[92] Bald Eagle courtship involves elaborate, spectacular calls and flight displays. The flight includes swoops, chases, and cartwheels, in which they fly high, lock talons, and free fall, separating just before hitting the ground.[43][93] Usually, a territory defended by a mature pair will be 1 to 2 km (0.62 to 1.2 mi) of waterside habitat.[7]

Compared to most other raptors which mostly nest in April or May, bald eagles are early breeders: nest building or reinforcing is often by mid-February, egg laying is often late February (sometimes during deep snow in the North), and incubation is usually mid-March and early May. Eggs hatch from mid April to early May, and the young fledge late June to early July.[7] The nest is the largest of any bird in North America; it is used repeatedly over many years and with new material added each year may eventually be as large as 4 meters (13 ft) deep, 2.5 meters (8.2 ft) across and weigh 1 metric ton (1.1 short tons);[2] one nest in Florida was found to be 6.1 meters (20 ft) deep, 2.9 meters (9.5 ft) across, and to weigh 3 short tons (2.7 t).[94] This nest is on record as the largest tree nest ever recorded for any animal.[95] Usually nests are used for under five years or so, as they either collapse in storms or break the branches supporting them by their sheer weight. However, one nest in the Midwest was occupied continuously for at least 34 years without interruption.[32] The nest is built out of branches, usually in large trees found near water. When breeding where there are no trees, the Bald Eagle will nest on the ground, as has been recorded largely in areas largely isolated from terrestrial predators, such as Amchitka Island in Alaska.[85] In Sonora, Mexico, eagles have been observed nesting on top of Hecho catcuses (Pachycereus pectinaboriginum).[96] Nests located on cliffs and rock pinnacles have been reported historically in California, Kansas, Nevada, New Mexico and Utah, but currently are only verified to occur only in Alaska and Arizona.[7] The eggs average about 73 millimeters (2.9 in) long, ranging from 58 to 85 mm (2.3 to 3.3 in), and have a breadth of 54 millimeters (2.1 in), ranging from 47 to 63 cm (19 to 25 in).[39][43] Eggs in Alaska averaged 130 g (4.6 oz) in mass, while in Saskatchewan they averaged 114.4 g (4.04 oz).[97][98] As with their ultimate body size, egg size tends to increase further away from the Equator.[43] Eagles produce between one and three eggs per year, two being typical. Rarely, four eggs have been found in nests but these may be exceptional cases of polygyny.[99] Eagles in captivity have been capable of producing up to seven eggs.[100] it is rare for all three chicks to successfully reach the fledging stage. The oldest chick often bear the advantage of larger size and louder voice, which tends to draw the parents attention towards them.[7] Occasionally, as is recorded in many large raptorial birds, the oldest sibling sometimes attacks and kills their younger sibling(s), especially early in the nesting period when their sizes are most different.[7] However, nearly half of known Bald Eagle produce two fledgings species (more rarely three), unlike in some other "eagle" species such as some in the Aquila genus, in which a second fledging is typically observed in less than 20% of nests despite two eggs being typically laid.[14] Both the male and female take turns incubating the eggs, but the female does most of the incubation. The parent not incubating will hunt for food or look for nesting material during this stage. For the first two to three weeks of the nestling period at least one adult is at the nest almost 100% of the time. After five to six weeks, the attendance of parents usually drops off considerably (with the parents often perching in trees nearby).[7] A young eaglet can gain up to 170 g (6.0 oz) a day, the fastest growth rate of any North American bird.[32] The young eaglets pick up and manipulate sticks, play tug of war with each other, practice holding things in their talons, and stretch and flap their wings. By eight weeks, the eaglets are strong enough to flap their wings, lift their feet off the nest platform, and rise up in the air.[32] The young fledge at anywhere from 8 to 14 weeks of age, though will remain close to the nest and attended to by their parents for a further 6 weeks. Juvenile eagles first start dispersing away from their parents about 8 weeks after they fledge. Variability in departure date related to effects of sex and hatching order on growth and development.[98] For the next four years, immature eagles wander widely in search of food until they attain adult plumage and are eligible to reproduce.[101] [10][11]Newly fledged juvenile===Longevity and mortality=== The average lifespan of Bald Eagles in the wild is around 20 years, with the oldest confirmed one having been 28 years of age.[4] In captivity, they often live somewhat longer. In one instance, a captive individual in New York lived for nearly 50 years. As with size, the average lifespan of an eagle population appears to be influenced by its location and access to prey.[102] As they are no longer heavily persecuted, adult mortality is quite low. In one study of Florida eagles, adult Bald Eagles reportedly had 100% annual survival rate.[8] In Prince William Sound in Alaska, adults had an annual survival rate of 88% even after the Exxon Valdez oil spill adversely effected eagles in the area.[103] Of 1,428 individuals from across the range necropsied by National Wildlife Health Center from 1963 to 1984, 329 (23%) eagles died from trauma, primarily impact with wires and vehicles; 309 (22%) died from gunshot; 158 (11%) died from poisoning; 130 (9%) died from electrocution; 68 (5%) died from trapping; 110 (8%) from emaciation; and 31 (2%) from disease; cause of death was undetermined in 293 (20%) of cases.[104] In this study, 68% of mortality was human-caused.[104] Today eagle-shooting is believed to be considerably reduced due to the species protected status.[105] In one case, an adult eagle investigating a Peregrine Falcon nest for prey items sustained a concussion from a swooping parent Peregrine, and ultimately died days later from it.[106] An early natural history video depicts a Cougar (Puma concolor) ambusing and killing an immature Bald Eagle feeding at a rabbit carcass is viewable online although this film may have been staged.[107]

Most non-human-related mortality involves nestlings or eggs. Around 50% of eagles survive their first year.[101] However, in the Chesapeake Bay area, 100% of 39 radio-tagged nestlings survived to their first year.[108] Occasionally, nestling or egg fatalities are due to nest collapses, starvation, sibling aggression or inclement weather. Another significant cause of egg and nestling mortality is predation. These have been verified to be predated by large gulls, corvids (including ravens, crows and magpies), Wolverines (Gulo gulo), hawks, owls, eagles, Bobcats (Lynx rufus), American black bears (Ursus americanus) and raccoons.[97][109][110][111][112][113][114] If food access is low, parental attendance at the nest may be lower because both parents may have to forage thus resulting in less protection.[14] Nestlings are usually exempt from predation by terrestrial carnivores that are poor tree-climbers, but Arctic Foxes (Vulpes lagopus) occasionally snatched nestlings from ground nests on Amchitka Island in Alaska before they were extirpated from the island.[85] The Bald Eagle will defend its nest fiercely from all comers and has even repelled attacks from bears, having been recorded knocking a black bear out of a tree when the latter tried to climb a tree holding nestlings.[115]

Relationship with humans

Population decline and recovery

Once a common sight in much of the continent, the Bald Eagle was severely affected in the mid-20th century by a variety of factors, among them the thinning of egg shells attributed to use of the pesticide DDT.[116] Bald Eagles, like many birds of prey, were especially affected by DDT due to biomagnification. DDT itself was not lethal to the adult bird, but it interfered with the bird's calcium metabolism, making the bird either sterile or unable to lay healthy eggs. Female eagles laid eggs that were too brittle to withstand the weight of a brooding adult, making it nearly impossible for the eggs to hatch.[26] It is estimated that in the early 18th century, the Bald Eagle population was 300,000–500,000,[117] but by the 1950s there were only 412 nesting pairs in the 48 contiguous states of the US. Other factors in Bald Eagle population reductions were a widespread loss of suitable habitat, as well as both legal and illegal shooting. In 1930 a New York City ornithologist wrote that in the state of Alaska in the previous 12 years approximately 70,000 Bald Eagles had been shot. Many of the hunters killed the Bald Eagles under the long-held beliefs that Bald Eagles grabbed young lambs and even children with their talons, yet the birds were innocent of most of these alleged acts of predation (lamb predation is rare, human predation is thought to be non-existent).[118] Later illegal shooting was described as "the leading cause of direct mortality in both adult and immature bald eagles," according to a 1978 report in the Endangered Species Technical Bulletin. In 1984, the National Wildlife Federation listed hunting, power-line electrocution, and collisions in flight as the leading causes of eagle deaths. Bald Eagles have also been killed by oil, lead, and mercury pollution, and by human and predator intrusion at nests.[119]

The species was first protected in the U.S. and Canada by the 1918 Migratory Bird Treaty, later extended to all of North America. The 1940 Bald Eagle Protection Act in the U.S., which protected the Bald Eagle and the Golden Eagle, prohibited commercial trapping and killing of the birds. The Bald Eagle was declared an endangered species in the U.S. in 1967, and amendments to the 1940 act between 1962 and 1972 further restricted commercial uses and increased penalties for violators. Perhaps most significant in the species' recovery, in 1972, DDT was banned from usage in the United States.[120] DDT was completely banned in Canada in 1989, though its use had been highly restricted since the late 1970s.[121] [12][13]First-yearWith regulations in place and DDT banned, the eagle population rebounded. The Bald Eagle can be found in growing concentrations throughout the United States and Canada, particularly near large bodies of water. In the early 1980s, the estimated total population was 100,000 individuals, with 110,000–115,000 by 1992;[2] the U.S. state with the largest resident population is Alaska, with about 40,000–50,000, with the next highest population the Canadian province of British Columbia with 20,000–30,000 in 1992.[2] Recent estimates indicate that the total population consists of approximately 200,000 eagles today.[122] For some time, the stronghold breeding population of Bald Eagles in the lower 48 states was in Florida, where over a thousand pairs have held on while populations in other states were significantly reduced by DDT use. Today, the contiguous state with the largest number of breeding pairs of eagles is Minnesota with an estimated 1,312 pairs, surpassing Florida's most recent count of 1,166 pairs. 23, or nearly half, of the 48 contiguous states now have at least 100 breeding pairs of Bald Eagles.[27]

The Bald Eagle was officially removed from the U.S. federal government's list of endangered species on July 12, 1995, by the U.S. Fish & Wildlife Service, when it was reclassified from "Endangered" to "Threatened." On July 6, 1999, a proposal was initiated "To Remove the Bald Eagle in the Lower 48 States From the List of Endangered and Threatened Wildlife." It was de-listed on June 28, 2007.[123] It has also been assigned a risk level of Least Concern category on the IUCN Red List.[1] In the Exxon Valdez Oil Spill of 1989 an estimated 247 were killed in Prince William Sound, though the local population returned to its pre-spill level by 1995.[4]

In captivity

Permits are required to keep Bald Eagles in captivity in the United States. Permits are primarily issued to public educational institutions, and the eagles which they show are permanently injured individuals which cannot be released to the wild. The facilities where eagles are kept must be equipped with adequate caging and facilities, as well as workers experienced in the handling and care of eagles. Bald Eagles cannot legally be kept for falconry in the United States. As a rule, the Bald Eagle is a poor choice for public shows, being timid, prone to becoming highly stressed, and unpredictable in nature. Native American Tribes can obtain a "Native American Religious Use" permit to keep non-releasable eagles as well. They use their naturally molted feathers for religious and cultural ceremonies. The Bald Eagle can be long-lived in captivity if well cared for, but does not breed well even under the best conditions.[124] In Canada, a license is required to keep Bald Eagles for falconry.

Known Bald Eagles

Bald eagles that appear in the Guardians of Ga'hoole series

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